Pop sensations Alvin, Simon and Theodore end up in the care of Dave Seville's twenty-something nephew Toby (Levi). The boys must put aside music superstardom to return to school, and are tasked with saving the school's music program by winning the $25,000 prize in a battle of the bands. But the Chipmunks unexpectedly meet their match in three singing chipmunks known as The Chipettes -– Brittany, Eleanor and Jeanette. Romantic and musical sparks are ignited when the Chipmunks and Chipettes square off.

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